The rounded long [ǭ] survived in Primitive Danian, as well as developing from primitive long [ā], but became unrounded [ō] in East Danian and the diphthong [ua] in Ossiriandic (PE19/25). It reappeared later in Ossiriandic as a development from the diphthong [au], but later still reduced to the unrounded long [ō].